Dr Adrian Willoughby

Senior Lecturer

Email: adrian.willoughby@monash.edu
Tel: +603 5514 6021
Office: Room 4-4-05

Profile Summary

Dr Adrian Willoughby is a Senior Lecturer in the School of Psychology at Monash University Malaysia.  He was awarded a BA (Hons) in Experimental Psychology from the University of Oxford and, in 2005, earned his PhD in Cognitive Psychology from the University of Michigan.  He uses behavioural, EEG and peripheral psychophysiology measures to study various cognitive processes, in particular attention, memory and executive control.

Prior to joining Monash University Malaysia in 2019, he was a Lecturer in the School of Psychology and Clinical Language Sciences at the University of Reading Malaysia in Johor.  Before moving to Malaysia, he worked as a Research Scientist in the Computer Science lab and Human Sleep lab at SRI International in California and has gained valuable research experience at institutions in both the USA and UK.

In addition to his research, Dr Willoughby has taught a variety of psychology courses at a number of institutions including the University of California, Berkeley and the University of California, Davis.


  • BA (Hons) in Experimental Psychology, University of Oxford
  • PhD in Cognitive Psychology, University of Michigan


Dr Willoughby’s current research interests centre on attention, memory and executive control – and the use of EEG, eye-tracking and peripheral psychophysiological methods to investigate them.

He is currently actively involved in research projects investigating multi-modal attention (how we divide our attention between visual and auditory input, and how processing of one type of information affects processing of the other), how our attention is affected by the contents of our working memory (what we are currently thinking about), mind wandering and multi-modal integration (how we combine information from different senses).


Willoughby A.R., de Zambotti M., Baker F.C., Colrain I.M. (In Press).  Evoked K-complexes and Altered Interaction Between the Central and Autonomic Nervous Systems during Sleep in Alcohol Use Disorder.  Alcohol.

Goldstone A., Willoughby A.R., de Zambotti M., Clark D.B., Sullivan E.V., Hasler B.P., Franzen P.L., Prouty D.E., Colrain I.M., Baker F.C. (2019).  Sleep spindle characteristics in adolescents.  Clinical Neurophysiology, 130(6), 893-902.

Goonawardena A.V., Morairty S.R., Orellana G.A., Willoughby A.R., Wallace T.L., and Kilduff T.S. (2019).  Electrophysiological characterization of sleep/wake, activity and the response to caffeine in adult cynomolgus macaques.  Neurobiology of Sleep and Circadian Rhythms, 6, 9-23.

Goldstone A., Willoughby A.R., de Zambotti M., Franzen P.L., Kwon D., Pohl K., Pfefferbaum A., Sullivan E., Muller-Oehring E., Prouty D., Hasler B., Clark D.B., Colrain I.M. and Baker F.C. (2018).  The mediating role of cortical thickness and gray matter volume on sleep slow-wave activity during adolescence.  Brain Structure and Function, 223(2), 669-685.

Willoughby A. R., de Zambotti M., Baker F.C. and Colrain I.M. (2015).  Partial K-complex recovery following short-term abstinence in individuals with alcohol use disorder.  Alcoholism: Clinical and Experimental Research, 39(8), 1417-24.

Cooke A., Kavussanu M., Gallicchio G., Willoughby A.R., McIntyre D. and Ring C. (2014).  Preparation for action: psychophysiological activity preceding a motor skill as a function of expertise, performance outcome, and psychological pressure.  Psychophysiology, 51(4), 374-84.

Kavussanu M., Willoughby A.R. and Ring C. (2012).  Moral Identity and Emotion in Athletes: Physiological Responses during Affective Picture Viewing.  Journal of Sport and Exercise Psychology, 34(6), 695-714.

Pratt N.L., Willoughby A.R. and Swick D. (2011).  Effects of working memory load on visual selective attention: behavioral and electrophysiological evidence.  Frontiers in Human Neuroscience, 5:57.

Gehring W.J. and Willoughby A.R. (2002).  The medial frontal cortex and the rapid processing of monetary gains and losses.  Science, 295, 2279-2282.

Research Grants


Other Achievements